The difficulty of answering this question goes to the issue of at point in time are we speaking? Moral behavior in the beginning may not at all resemble moral thinking today. Early man believed that human sacrifaces to the gods were a good thing. I suspect that since human beings are superstitious gods in some form have existed from the beginning and hence some form of religion. I suspect that through man's evolutionary process moral behavior and religion merged. for instance a definition of moral behavior today is "doing the right thing when no one is watching." Primative religious men believed human sacrifaces in private were for the public good in appeasing the gods. Hedondism was acceptable behavior and not thought immoral.
Eskimoes believed it was good manners to share their wives with vistors.
so if the definition of religion and moral behavior is very liberally applied then my view would be religion first owing to man's superstitious nature and then moral behavior as man began to be domesticated
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